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Polyploidy and subsequent post-polyploid diploidization (PPD) are key drivers of plant genome evolution, yet their contributions to evolutionary success remain debated. Here, we analyze the Malvaceae family as an exemplary system for elucidating the evolutionary role of polyploidy and PPD in angiosperms, leveraging 11 high-quality chromosome-scale genomes from all nine subfamilies, including newly sequenced, near telomere-to-telomere assemblies from four of these subfamilies. Our findings reveal a complex reticulate paleoallopolyploidy history early in the diversification of the Malvadendrina clade, characterized by multiple rounds of species radiation punctuated by ancient allotetraploidization (Mal-β) and allodecaploidization (Mal-α) events around the Cretaceous–Paleogene (K–Pg) boundary. We further reconstruct the evolutionary dynamics of PPD and find a strong correlation between dysploidy rate and taxonomic richness of the paleopolyploid subfamilies (R^2 ≥ 0.90, P < 1e-4), supporting the “polyploidy for survival and PPD for success” hypothesis. Overall, our study provides a comprehensive reconstruction of the evolutionary history of the Malvaceae and underscores the crucial role of polyploidy–dysploidy waves in shaping plant biodiversity. 

Abstract Whistler mode waves scatter energetic electrons, causing them to precipitate into the Earth's atmosphere. While the interactions between whistler mode waves and electrons are well understood, the global distribution of electron precipitation driven by whistler mode waves needs futher investigations. We present a two‐stage method, integrating neural networks and quasi‐linear theory, to simulate global electron precipitation driven by whistler mode waves. By applying this approach to the 17 March 2013 geomagnetic storm event, we reproduce the rapidly varying precipitation pattern over various phases of the storm. Then we validate our simulation results with POES/MetOp satellite observations. The precipitation pattern is consistent between simulations and observations, suggesting that most of the observed electron precipitation can be attributed to scattering by whistler mode waves. Our results indicate that chorus waves drive electron precipitation over the premidnight‐to‐afternoon sector during the storm main phase, with simulated peak energy fluxes of 20 erg/cm²/s and characteristic energies of 10–50 keV. During the recovery phase, plume hiss in the afternoon sector can have a comparable or stronger effect than chorus, with peak fluxes of ∼1 erg/cm²/s and characteristic energies between 10 and 200 keV. This study highlights the importance of integrating physics‐based and deep learning approaches to model the complex dynamics of electron precipitation driven by whistler mode waves. 

Auroral precipitation is the second major energy source after solar irradiation that ionizes the Earth’s upper atmosphere. Diffuse electron aurora caused by wave-particle interaction in the inner magnetosphere (L < 8) takes over 60% of total auroral energy flux, strongly contributing to the ionospheric conductance and thus to the ionosphere-thermosphere dynamics. This paper quantifies the impact of chorus waves on the diffuse aurora and the ionospheric conductance during quiet, medium, and strong geomagnetic activities, parameterized by AE <100, 100 < AE < 300, and AE > 300, respectively. Using chorus wave statistics and inner-magnetosphere plasma conditions from Timed History Events and Macroscale Interactions during Substorms (THEMIS) observations, we directly derive the energy spectrum of diffuse electron precipitation under quasi-linear theory. We then calculate the height-integrated conductance from the wave-driven aurora spectrum using the electron impact ionization model of Fang et al. (Geophys. Res. Lett., 2010, 37) and the MSIS atmosphere model. By utilizing Fang’s ionization model, the US Naval Research Laboratory Mass Spectrometer and Incoherent Scattar Radar (NRLMSISE-00) model from 2000s for the neutral atmosphere components, and the University of California, Los Angeles (UCLA) Full Diffusion Code, we improve upon the standard generalization of Maxwellian diffuse electron precipitation patterns and their resulting ionosphere conductance. Our study of global auroral precipitation and ionospheric conductance from chorus wave statistics is the first statistical model of its kind. We show that the total electron flux and conductance pattern from our results agree with those of Ovation Prime model over the pre-midnight to post-dawn sector as geomagnetic activity increases. Our study examines the relative contributions of upper band chorus (UBC) and lower band chorus wave (LBC) driven conductance in the ionosphere. We found LBC waves drove diffuse electron precipitation significantly more than UBC waves, however it is possible that THEMIS data may have underestimated the upper chorus band wave observations for magnetic latitudes below 65 $°$. 

Abstract We show that two semi‐infinite positive temperature polymers coalesce on the scale predicted by KPZ (Kardar–Parisi–Zhang) universality. The two polymer paths have the same asymptotic direction and evolve in the same environment, independently until coalescence. If they start at distance apart, their coalescence occurs on the scale . It follows that the total variation distance of two semi‐infinite polymer measures decays on this same scale. Our results are upper and lower bounds on probabilities and expectations that match, up to constant factors and occasional logarithmic corrections. Our proofs are done in the context of the solvable inverse‐gamma polymer model, but without appeal to integrable probability. With minor modifications, our proofs give also bounds on transversal fluctuations of the polymer path. As the free energy of a directed polymer is a discretization of a stochastically forced viscous Hamilton–Jacobi equation, our results suggest that the hyperbolicity phenomenon of such equations obeys the KPZ exponent. 

We analyze the properties of relativistic (>700 keV) electron precipitation (REP) events measured by the low-Earth-orbit (LEO) POES/MetOp constellation of spacecraft from 2012 through 2023. Leveraging the different profiles of REP observed at LEO, we associate each event with its possible driver: waves or field line curvature scattering (FLCS). While waves typically precipitate electrons in a localized radial region within the outer radiation belt, FLCS drives energy-dependent precipitation at the edge of the belt. Wave-driven REP is detected at any MLT sector and L shell, with FLCS-driven REP occurring only over the nightside–a region where field line stretching is frequent. Wave-driven REP is broader in radial extent on the dayside and accompanied by proton precipitation over 03–23 MLT, either isolated or without a clear energy-dependent pattern, possibly implying that electromagnetic ion cyclotron (EMIC) waves are the primary driver. Across midnight, both wave-driven and FLCS-driven REP occur poleward of the proton isotropic boundary. On average, waves precipitate a higher flux of >700 keV electrons than FLCS. Both contribute to energy deposition into the atmosphere, estimated of a few MW. REP is more associated with substorm activity than storms, with FLCS-driven REP and wave-driven REP at low L shells occurring most often during strong activity (SML* < −600 nT). A preliminary analysis of the Solar Wind (SW) properties before the observed REP indicates a more sustained (∼5 h) dayside reconnection for FLCS-driven REP than for wave-driven REP (∼3 h). The magnetosphere appears more compressed during wave-driven REP, while FLCS-driven REP is associated with a faster SW of lower density. These findings are useful not only to quantify the contribution of >700 keV precipitation to the atmosphere but also to shed light on the typical properties of wave-driven vs FLCS-driven precipitation which can be assimilated into physics-based and/or predictive radiation belt models. In addition, the dataset of ∼9,400 REP events is made available to the community to enable future work. 

Abstract Whistler‐mode waves are commonly observed within the lunar environment, while their variations during Interplanetary (IP) shocks are not fully understood yet. In this paper, we analyze two IP shock events observed by Acceleration, Reconnection, Turbulence and Electrodynamics of the Moons Interaction with the Sun (ARTEMIS) satellites while the Moon was exposed to the solar wind. In the first event, ARTEMIS detected whistler‐mode wave intensification, accompanied by sharply increased hot electron flux and anisotropy across the shock ramp. The potential reflection or backscattering of electrons by the lunar crustal magnetic field is found to be favorable for whistler‐mode wave intensification. In the second event, a magnetic field line rotation around the shock region was observed and correlated with whistler‐mode wave intensification. The wave growth rates calculated using linear theory agree well with the observed wave spectra. Our study highlights the significance of magnetic field variations and anisotropic hot electron distributions in generating whistler‐mode waves in the lunar plasma environment following IP shock arrivals. 

Abstract We statistically evaluate the global distribution and energy spectrum of electron precipitation at low‐Earth‐orbit, using unprecedented pitch‐angle and energy resolved data from the Electron Losses and Fields INvestigation CubeSats. Our statistical results indicate that during active conditions, the ∼63 keV electron precipitation ratio peaks atL > 6 at midnight, whereas the spatial distribution of precipitating energy flux peaks between the dawn and noon sectors. ∼1 MeV electron precipitation ratio peaks near midnight atL > ∼6 but is enhanced near dusk during active times. The energy spectrum of the precipitation ratio shows reversal points indicating energy dispersion as a function ofLshell in both the slot region and atL > ∼6, consistent with hiss‐driven precipitation and current sheet scattering, respectively. Our findings provide accurate quantification of electron precipitation at various energies in a broad region of the Earth's magnetosphere, which is critical for magnetosphere‐ionosphere coupling. 

In this study, we present simultaneous multi-point observations of magnetospheric oscillations on a time scale of tens of minutes (forced-breathing mode) and modulated whistler-mode chorus waves, associated with concurrent energetic electron precipitation observed through enhanced BARREL X-rays. Similar fluctuations are observed in X-ray signatures and the compressional component of magnetic oscillations, spanning from ∼9 to 12 h in MLT and 5 to 11 inLshell. Such magnetospheric oscillations covering an extensive region in the pre-noon sector have been suggested to play a potential role in precipitating energetic electrons by either wave scattering or loss cone modulation, showing a high correlation with the enhancement in X-rays. In this event, the correlation coefficients between chorus waves (smoothed over 8 min), ambient magnetic field oscillations and X-rays are high. We perform an in-depth quasi-linear modeling analysis to evaluate the role of magnetic field oscillations in modulating energetic electron precipitation in the Earth’s magnetosphere through modulating whistler-mode chorus wave amplitude, resonance condition between chorus waves and electrons, as well as loss cone size. Model results further show that the modulation of chorus wave amplitude plays a dominant role in modulating the electron precipitation. However, the effect of the modulation in the resonant energy between chorus waves and energetic electrons due to the background magnetic field oscillations cannot be neglected. The bounce loss cone modulation, affected by the magnetic oscillations, has little influence on the electron precipitation modulation. Our results show that the low frequency magnetospheric oscillations could play a significant role in modulating the electron precipitation through modulating chorus wave intensity and the resonant energy between chorus waves and electron.